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Doubtful those listed are to blame, though it is most likely from bullying behavior from somewhere.
Can you tell us more about the system and inhabitants?
29g BioCube with 2 Ocellaris Clownfish, 2 Banggai Cardinalfish, and the Blue Fin Goby. Inverts - 3 Dwarf Red Tip Hermit Crabs, 4 Scarlet Reef Hermit Crabs, 3 Dwarf Yellow Tip Hermit Crabs, the Shrimp mentioned above, 3 Mexican Turbo Snails, 3 Cerith Snails, 2 Super Tongan Nassarius Snails, and Fighting Conch. That's the full stock.