Just wanted to make a small point. JMcpeak, said Quote " The anaerobic bacteria that form in the mud utilise Nitrate, just the same as nitrosamonas/nitrospira utilise ammonia and nitrite ".
That's not quite right. Both bacterial processes convert toxins into products that are less toxic, or not toxic, but the way they do it is different. The Aerobic BB in FW filters consume ammonia and nitrite, the end product is nitrate, which is still a toxin that has to be managed, typically by water changes and/or plantings as well.
The anaerobic bacteria you get in a DSB, more usual in salt tanks, go a step further, converting nitrate into Nitrogen gas, which dissipates into the air. The gas is entirely non toxic, needing no further management.
There's a downside to anaerobic bacteria, because if they get out of hand they can also produce other gasses, like hydrogen sulphide gas, which is pretty nasty and can kill fish or inverts quite easily.
Our fresh water BB are actually nitrifiers, & the anaerobes are DEnitrifiers in a DSB.