Thant is an interesting line of thought. However I have a few problems with it. First of a jaguar Cichlid (Im assuming you are referring to Parachromis managuensis) is inherently an aggressive and territorial species. It will give most things you put in with it a hard time and that is not because it is trying to breed, it is because it is very intolerant of another fish within an enclosed space. A caged animal is almost 100% of the time more aggressive than one out in the Wild. A single juvenile Jag will be aggressive like a pair of Breeding adults. If juvenile who has no drive to breed yet is still acting aggressively this would pretty much prove it is not just a breeding behavior. I believe this pretty much debunks the theory that they are aggressive only because they are attempting to breed.
Many Fish in the wild also spawn regularly, some almost constantly like many spawn scatters including rainbows, tetras, barbs, minnows, Etc. Many cichlids are no exception. Many rift lake species are almost always either in the process of raising fry or laying eggs, SA and CA species also breed quite regularly with their partner and can raise many spawns successively after the other. The food that these animals often consume in the wild is far superior to our processed foods in terms of conditioning for spawning. Algae eaters and grazers have an almost infinite supply of bio-film and the like. Characins, including tetras, silver Dollars, Pencilfish, and Piranhas have a vast supply of insect larvae, other small fish fry, infusoria, etc to feed on constantly throughout the day, Many breeders try to emulate this in order to stimulate courtship. In these species. Also, addressing your last point, packing a tank full of fish does stress a fish out. Many fish can be outcompeted for food by boisterous tank mates and may species become increasingly shy with every tank mate added (Think CPDs and Micro fish)
I disagree with some of that and agree with some as well.
I will mainly focus on larger cichlids and other larger predators as that was what i was originally talking about with my argument of them always being in breeding mode. These fish are also usually the cause of threads such as this one, because the bigger the fish the more difference in opinions on what tank they need. I understand that many others arent like this. I merely used an example that describes a lot of fishes.
Sure, food in the wild is better for them, they lived for thousands if not millions of years eating it. But they don't get food every day in mist cases. Sometimes they don't even get it every single week. Its generally at certain times during the year when they spawn, which is usually in springtime for most fish. Im not exactly sure when it is in a more stable climate though.
Example: the common types of Sunfish you catch at a lake, Bluegills and Pumpkinseeds etc. These only usually spawn during spring. Plants grow then, other fish eat off of plants and other tiny organisms that need warm water, which gives these fish more food. They are then able to condition themselves for spawning. After they downsize the population of any fishes they can eat, it becomes harder for them to spawn. Females of any species need a lot of nutrients to create eggs, and any live-bearing animals need even more. The males are able to spawn much more often than females, but still need to be healthy.
Also, in regards to the Jaguar, and yes i mean the Managuensis; they are trying to breed at a young age. You can find examples of 2-3 inch Jags breeding, although most fry didn't survive. As soon as they are big enough to produce the things they need, fertile eggs and sperm, they will breed. This happens very early. However, in the wild it is basically impossible for these fish to breed at this size. Because there is no way for them to get the nutritional value to be in a condition where that can survive breeding. They simply cannot get enough food in their bodies to breed.
So yes, as soon as you see them getting aggressive they are trying to breed. They defend territories mainly for breeding purposes.
Also, caged animals arent as aggressive in all cases as wild animals. This boils down in a few ways.
Generally they arent as predatory. They arent being fed live creatures as a staple, like they would in the wild. Predators are generally more aggressive as they have to compete harder for food.
Also, in a sense of being around humans, they arent really being more aggressive. They just arent scared anymore. A Dovii that is 20+ inches long in aquarium isn't scared to swim towards you and take a chomp, but in the wild they would make a quick escape.
And in ways, yes overstocking does stress fish. But, as for this specific argument I'm sticking to the bug predators, i kind of disagree. They arent really out-competed. They are usually able to pack enough punch to dive in and get a bite or two of food and get out, and repeat. In the wild they have to deal with this in ways as well. My reasoning is this: since a lot of prey fish school, the predators are usually attracted to these giant masses of baitfish. This is and easier meal for them that can really help them out by filling them up for a while.
The big fish just dive right in, and eat a lot of those smaller fish. The predators that are smaller but still wanting a meal have to take a chance; they can go in and get a good meal and maybe get bitten by a bigger fish, or stay out and stay safe. They still go for the food, even though the big fish are in a bit of a frenzy and might eat the smaller predators. They usually can grab a bite and swim away safely.
Its usually the same in the aquarium. With a lot of food, the fish get into an excited state. They are focused on food and not chasing down their opponents. We also usually keep fish close enough in size to not eat each other.
Does that make any sense? Again, im focusing on the larger types of predators, think Convict and larger. These are the fish that have to get filled with nutrients to breed, which is hard in the wild due to a supply of food that isn't a constant for them, but is in our tanks.
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